P-22 was believed to be 12 years old, longer-lived than most wild male mountain lions.
was confirmed by a scan that revealed injuries to his head and torso, wildlife officials said.
P-22 was born in the western Santa Monica Mountains, and he was partly famous for apparently crossing two heavily traveled freeways on his trek east to the 4,200-acre park in the urbanized eastern end of the mountain range. P-22 was outfitted with a tracking collar in 2012. A year later his celebrity was solidified when he appeared in afeature with an iconic photo of the big cat on an LA hillside with the Hollywood sign in the background.
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