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Eocene giant ants, Arctic intercontinental dispersal, and hyperthermals revisited: discovery of fossil Titanomyrma (Hymenoptera: Formicidae: Formiciinae) in the cool uplands of British Columbia, Canada | The Canadian Entomologist | Cambridge Core

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Eocene giant ants, Arctic intercontinental dispersal, and hyperthermals revisited: discovery of fossil Titanomyrma (Hymenoptera: Formicidae: Formiciinae) in the cool uplands of British Columbia, Canada | The Canadian Entomologist | Cambridge Core
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'Giant' ant fossil raises questions about ancient Arctic migrations sfu

: joins close to m-cu], cell 1Rs evenly widens distally [narrows distad 2r], cell 1M about as high as wide [about half as high as wide], cu-a joins M+Cu at approximately half its length [joins very close to 1-M, Cu].

Hind wings missing. Legs: portions preserved short, stout. Waist: single segmented, petiole lacking anterior peduncle, otherwise indistinctly preserved. Partial propodeal spiracle preserved, slit-like, long. Gaster: only the first segment preserved. Beverly Burlingame , 12.vi.2021.Head: missing antennae; head length, ∼ 5.1 mm; head width, ∼ 6.5 mm; eye length, ∼ 1.1 mm, head capsule about 2/5 alitrunk length , with rounded posterior margins, not lobed; eyes large, oval, maximum length oriented anterodorsally, located at cephalic mid-length; large, triangular mandibles, about half head length, with approximately 8–10 coarse teeth. Alitrunk: somewhat distorted, indistinctly preserved; wings not preserved; right legs partially preserved, short, stout, left legs absent. Spiracles slit-like, long; propodeal spiracles robust, widest; petiole spiracles smaller, thinner; one gaster spiracle present, poorly preserved. Waist: single segmented, petiole lacking anterior peduncle, apparently about 5 mm wide, otherwise indistinct. Gaster: small, indistinct portion preserved. Paul Buchheim , 7.viii.2002.Head missing. Alitrunk: well preserved basally, faintly in anterior two thirds; legs short, stout; femur, tibia of the left mesothoracic and metathoracic legs, the right metathoracic leg, the femur of right mesothoracic leg, and tibia of the left prothoracic present. Wings not preserved. Spiracles slit-like, long; propodeal spiracle robust, widest; petiole spiracle shortest; petiole, A3, A4 spiracles narrower; A3, A4 spiracles long. Waist: single segmented, petiole about 4.5 mm wide, length indeterminate, apparently lacking anterior peduncle. Gaster: A3, about 9 mm wide, length indeterminate; A4, 14 mm wide, about 4.5–5.0 mm long, missing posterior to this. Paul Buchheim , 25.vii.2000.The new Allenby specimen is the first occurrence of Formiciinae in a known microthermal climate. Its implications for the relationships between queen size, climate, and trans-Arctic dispersal are complicated, however, by uncertainty of its size in life, as its sole fossil has been distorted by geologic shear forces during diagenesis, a common occurrence in the exposure where it was found (see Archibald and Cannings

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